CUPBChina University of Petroleum, Beijing
CUPBCornell University Program Board
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2012), [dagger]uralica possesses a common stem composed of M (green) and CuA (orange), a branched M (into MA and MP), a distinct free part of CuA, and CuP (purple) giving rise to CuPa (purple) and CuPb, a CuPa giving rise to CuPaa (blue) and CuPa[beta] (red), and a fusion of CuA with CuPa[alpha] resulting into a composite stem.
2G-H); RA simple; MA simple; CuA + CuPa[alpha] with two branches (i.e., CuPaa simple; in the right forewing CuPa[alpha] is probably forked with the two resulting branches re-uniting into a single stem); CuA, CuPa[alpha] , CuPa[beta] , and AA1 and other anal veins reaching wing margin in a very distal position; intercalary veins occur in areas between CuPa[beta] and CuPb, CuPb and AA1, and between anal veins.
Although the basal stem of CuP can be identified from the wing base (it gives rise to CuPa[beta] and CuPb, and is located anterior to AA1; Fig.
mexicanus to be distinguished from that of Palaeozoic stem-orthopterans are: RA simple (possibly a consequence of abbreviation); lack of strong inflexion point of the wing outline opposite the end of CuPb; CuA + CuPa[alpha] fusing with CuPa[beta] (near posterior wing margin).
Female forewing: 45 mm long, width 18 mm, various cross-veins filling the veinlets between ScA and anterior margin; RP branched opposite to the second branch of RA, RA pectinate with 7 branches, RP with at least 5 branches; MP parallel with MA, but not sigmoidal; CuA nearly the same length as free part of M; CuA+ CuPa[alpha] with 5 branches, first posterior branch emitted slightly before the fusion of CuA and CuPa[alpha]; CuPa[beta] oblique and straight; CuPb slightly curved in basal part.
Forewing oval, anterior margin arched, R forked very distally, M + CuA curved towards R, area between origin of RP and M distinctly broad, 1A bowed towards posterior margin and distant to CuPb.
It's very similar to some Tuphellid insects which are only found in the Triassic and to Haglinae; however this taxon shares with Aboilinae a well-developed ScA which, cutting the branches of ScP, MA and MP, is simple and without any anastomosis with RP; a series of straight cross-veins fill up the proximal part of CuPb and CuPa[beta].
It is similar to Circulaboilus aureus Li, Ren & Wang, 2007, (Cyrtophyllitinae) but we can separate it by the following characters: M + CuA strongly curved towards R, R forked very distally at about 2/3 of the wing length, RP basally curved, 2A bowed towards posterior margin and distant to CuPb. It is also similar to Barchaboilus mongolicus Gorochov, 1988 and Tettaboilus pulcher Gorochov, 1988, but they are different in their divergence of R, long ScA, more widened area between R and M + CuA and strongly oblique CuPa[beta].
Diagnosis.--Forewings: basal to the fusion of CuA with CuPa, area between ScA and ScP with a dense network of strong crossveins; R branched basally; RP simple for a long distance; RP and M branched; CuA (diverging from M + CuA) long; CuA + CuPa with numerous branches, without weak posterior veinlets vanishing in the area between CuA + CuPa and CuPb; crossvein network loose.
The condition of CuPb in hind wings (branched or simple) is unknown in most Palaeozoic Archaeorthoptera.
In summary, the combination of a simple CuPa, a common stem MP + CuA (diverging from M + CuA), a brief connection of the anterior branch of MA with RP, and of a branched CuPb, picture a hind wing that challenges assignment to any of the known Palaeozoic Archaeorthoptera.