INOSCIntegrated Network Operations and Security Center
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Specifically, the two INOSCs have purview over geographic regions (INOSC East and INOSC West); they configure and operate core services across the base networks in their domain, responsible for most base boundary protection and network security devices (the INOSC runs most network-defense software tools and devices even though they might be physically present at the local base).
For historical reasons, each major command generally has an INOSC detachment handling the more routine aspects of core network services across the command.
INOSC responsibilities include active directory management, mail relay, firewall management, web proxy services, gateway management and other operations associated with Air Force networks.