1 nests (75%) per year at PNMB over the same time period.
As a result, the geographic distribution of leatherback nesting in Costa Rica has been reduced to the Santa Elena and Nicoya peninsulas and is currently constricted to around an area about ~50 km north and south of PNMB.
However, the number of nesting turtles has declined to critically low levels, with only 26 turtles identified nesting at PNMB in both 20142015 and 2015-2016 (historically lowest levels) and <5 turtles per beach per season on secondary beaches in the last five years.
As expected, a similar declining trend to the one documented at PNMB was observed at Naranjo and Caletas, the two longest running projects on secondary beaches (Fig.
Beach exchange was previously reported to occur within index beaches of PNMB (Santidrian-Tomillo et al.