On the ventral surface, where the veins are most visible, VF1, VF3, VH1, and VH3 roughly measured the length of the wing; VF2, VF4, VH2, and VH4 measured width.
In all, 29 of the 45 traits were selected to construct the G-matrix used in one matrix test (EYE1, EYE2, EYE3, EYE4, EYES, EYE6, EYE7, EYE8, EYE9, EYE10, EYE11, DIAM3, DIAM4, DIAM11, EP1, EP2, EP3, EP6, EP8, PFE1, PFE2, CSS1, CSS2, DF1, DF2, DF3, VF4, VH1, VH4).